Journal of Conchology 44/4

CONChOlOgICAl SOCIETY Of grEAT BrITAIN ANd IrElANd www . CONCHSOC . ORg

h on . P residenT Tom Walker, 38 Redlands Rd, Reading,

h on . M eMBershiP l iason o FFicer Pat Robbins, 125 East Lane, West Horsley, Leatherhead, Surrey KT23 6LJ membership@conchsoc.org A cTinG h on . P roGraMMe s ecreTary Martin Willing, 14 Goodwood Close, Midhurst, programme@conchsoc.org h on . c onserVaTion o FFicer Mags Cousins, conservation@conchsoc.org h on . P uBlicaTions s erVices M anaGer (for back issues of the Journal and Mollusc World) Tom Walker, sales@conchsoc.org Journal of Conchology Editorial Board h on . e diTor – Anna M Holmes Associate Editors West Sussex, GU29 9JG

h on . M arine r ecorder Simon Taylor, Fiddlesticks, 44 Strawberry Lane, Tolleshunt Knights, Essex CO5 0RX

Berkshire, RG1 5HD president@conchsoc.org

h on . n on - Marine r ecorder Ben Rowson, Amgueddfa Cymru – National Museum of Wales, Dept. Natural Sciences, Cathays Park, Cardiff CF10 3NP

h on . G eneral s ecreTary Rosemary Hill, 447b Wokingham Road, Earley, Reading, RG6 7EL h on . T reasurer Brian Goodwin, 44 Amber Crescent, Walton, Chesterfield, Derbyshire, S40 3DH treasurer@conchsoc.org

h on . e diTor M ollusc W orld Peter Topley, The Rectory, 8 Rectory Close, Clifton, Shefford, Bedfordshire SG17 5EL

Robert Cameron (UK) Barna P Gergely (Hungary) David Holyoak (Portugal) Michal Horsak (Czech Republic) Manuel Malaquias (Portugal)

Daniella R Bar Yosef Mayer (Israel) Julia Piza (Argentina)

Julia Sigwart (USA/Ireland) Martin Zuschin (Austria) Ben Rowson (UK) Diego Zelaya (Argentina)

Dinarzarde Raheem (UK) Ira Richling (Germany) Carmen Salas (Spain)

SuBSCrIPTION ChArgES ANd PuBlICATIONS INfOrMATION i ndiVidual MeMBershiP UK addresses: £33 per annum; Rest of Europe (including Republic of Ireland): £41 per annum; Rest of the World: £50 per annum. F aMily MeMBershiP UK addresses: £35 per annum; Rest of Europe (including Republic of Ireland): £43 per annum; Rest of the World: £52 per annum. s TudenT ’ s MeMBershiP UK addresses: £15 per annum; Rest of Europe (including Republic of Ireland): £23 per annum; Rest of the World: £32 per annum. i nsTiTuTional suBscriPTion UK addresses: £47 per annum; Rest of Europe (including Republic of Ireland): £54 per annum; Rest of the World: £62 per annum. *Overseas members are reminded that all monies due to the Society are payable in sterling

P uBlicaTions Members receive: The Journal of Conchology (usually two numbers a year) Mollusc World (three a year)

For subscriptions, please contact: Carolyn Postgate at 14 St Barnabas Court, Cambridge CB1 2BZ

For back-numbers of these publications and special numbers please apply to: Tom Walker, sales@conchsoc.org

© Conchological Society of Great Britain and Ireland Typeset and printed in the United Kingdom by Henry Ling Limited, at the Dorset Press, Dorchester DT1 1HD Cover illustration: Trivia monacha photograph by Ian F. Smith https://www.researchgate.net/profile/Ian_Smith19/publications Date of publication: 7th October 2022